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Gar W. Rothwell

Gar Rothwell

Professor (Courtesy)
Ph.D., University of Alberta, 1973


Office/Lab: Cordley 1071/1090
Phone: 541-737-5252



Research Area

Studies of fossil and living land plants are directed toward a fuller understanding of phylogeny and evolution. These are explored using data from morphological, anatomical, ultrastructural, developmental, and molecular characters. Evaluations of ontogeny, reproductive biology, and organismal interactions are emphasized in interpreting development of the modern flora within the context of evolutionary ecology.

Jurassic conifer Longitudinal section of the Jurassic conifer Pararaucaria patigonica Wieland, which provides the first evidence for internal anatomy of seed cones assignable to the extinct family Cheirolepidiaceae (see Escapa et al., 2012 for details).


Related Links

Ohio University web site

Gymnosperms on the Tree of Life: Resolving the Phylogeny of Seed Plants

Paleobotany in Antarctica

Midcontinent Paleobotanical Colloquium

International Organisation of Paleobotany

Research Group Members

Graduate Students

Brian Atkinson (PhD due 2016)

Selected Publications

Nadon, G.C., A.M.F. Tomescu, L. Pratt, and G.W. Rothwell. 2013.  Differentiating among depositional environments with C/S. Data: An example from the Late Ordovician, south-central Pennsylvania, USA. Paleoecology, Paleogeography, Paleoclimatology, in revision.

Rothwell, G.W. and R.A. Stockey. 2013. Evolution and phylogeny of gnetophytes: evidence from the anatomically preserved seed cone Protoephedrites eamesii gen. et sp. nov. International Journal of Plant Sciences, in revision.

Serbet, R., B. Bomfleur and G.W. Rothwell. 2013.  Cunninghamia taylorii sp. nov. a structurally preserved cupressaceous conifer from the Upper Cretaceous (Campanian) Horseshoe Canyon Formation of Western North America. International Journal of Plant Sciences, in press.

Stockey, R.A., G.L. Hoffman, and G.W. Rothwell. 2013. Paleobotany and paleoecology of Gao Mine, a Late Paleocene fossil locality near Red Deer, Alberta, Canada. Canadian Journal of Earth Sciences, in press.

Stockey, R.A. and G.W. Rothwell. 2013. Pararaucaria carrii sp. nov., anatomically preserved evidence for the conifer family Cheirolepidiaceae in the Northern Hemisphere.  International Journal of Plant Sciences, in press.

Escapa, I.H., N.R. Cuneo, G.W. Rothwell and R.A. Stockey, 2013. Pararaucaria delfueyoi from the Late Jurassic Cañadón Calcáreo Formation, Chubut, Argentina: insights into the evolution of Cheirolepidiaceae.  International Journal of Plant Sciences, in press.

Ryberg, P.E., R.A. Stockey, J. Hilton, G. Mapes, J.B. Riding and G.W. Rothwell. 2012.  Reconsidering relationships among stem and crown group Pinaceae: Oldest record of the genus Pinus from the Early Cretaceous of Yorkshire, UK. International Journal of Plant Sciences, in press.

Rothwell, G.W., G. Mapes, R.A. Stockey and J. Hilton. 2012. The seed cone Eathiestrobus gen. nov.; fossil evidence for a Jurassic origin of Pinaceae.  American Journal of Botany 99: 708-720.

Escapa, I.H., G.W. Rothwell, R.A. Stockey, and N.R. Cuneo. 2012. Seed cone anatomy of Cheirolepidiaceae (Coniferales): Reinterpreting Pararaucaria patagonica Wieland.  American Journal of Botany 99:1058-1068.

Dunn, M. and G.W. Rothwell. 2012. Phenotypic placticity of the hydrasperman seed fern Tetrastichia bupatides Gordon. International Journal of Plant Sciences 157:823-834.

Hillier, K.G. and G.W. Rothwell. 2012. Senftenbergia oregonensis (Arnold) Comb. Nov., (Filicales); delimiting relationships among Paleozoic Tedeleaceae. International Journal of Plant Sciences 173: 549-557.

Sanders. H., G.W. Rothwell and S.W. Wyatt. 2011. Parallel Evolution of Auxin Regulation in Rooting Systems. Plant Systematics and Evolution 291: 221-225.

Klymiuk, A., G.W. Rothwell and R.A. Stockey. 2011. The first organismal concept for an extinct species of Pinaceae: Pinus arnoldii Miller. International Journal of Plant Sciences 172: 294-313.

Rothwell, G.W., R. A. Stockey, G. Mapes, and J. Hilton. 2011. Structure and relationships of the Jurassic conifer seed cone Hughmillerites juddii gen. et comb. nov. : implications for the early evolution of Cupressaceae.  Review of Palaeobotany and Palynology164: 45-59.

Jud, N.A., G.W. Rothwell and R.A. Stockey. 2010. Paleoecological and phylogenetic implications of Saxicaulis meckertii gen. et sp. nov.: a bennettitalean stem from the Upper Cretaceous of western North America. International Journal of Plant Sciences. 171: 915-925.

Stevens, L.G., J. Hilton, A.R. Rees, G.W. Rothwell and R.M. Bateman. 2010. Systematics, phylogenetics, and reproductive biology of Flemingites arcuatus sp. nov., an exceptionally preserved and partly reconstructed Carboniferous arborescent lycopsid. International Journal of Plant Sciences 171: 783-808.

Rothwell, G.W. and R.A. Stockey. 2010. Independent evolution of seed enclosure in Bennettitales: Evidence from the anatomically preserved cone Foxeoidea connatum gen. et sp. nov.  Pp. 51-66. In C.T. Gee [ed.], Plants in Mesozoic Time: Innovations, Phylogeny, Ecosystems (Ted Delevoryas festschrift).  Indiana University Press, Bloomington and Indianapolis.


Stewart, W.N. and G.W. Rothwell. 1993. Paleobotany and the evolution of plants. Cambridge University Press, 521 pp.